Penn State Altoona 1 article published in JoVE Neuroscience Assessing Spatial Learning and Memory in Small Squamate Reptiles Lara D. LaDage1, Tracie E. Cobb Irvin1, Victoria A. Gould1 1Division of Mathematics and Natural Sciences, Penn State Altoona This paper describes a modification of the Barnes maze, a standard rodent paradigm used to assess spatial memory and learning, for use in small squamate reptiles.