Name: photosystem i and nadph formation
Uploaded: 2019-03-11T09:21:12.000Z
Duration: 1 min 27 s
Channel: JoVE

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Earth’s Systems

HS-ESS2-6

Photosystem I

Подобно фотосистеме II, в фотосистеме I, фотоны также поглощаются молекулами пигмента в светособирающем комплексе, чтобы связать энергию с центром реакции и после окисления, высокоэнергетический электрон передаётся к первичному электронному акцептору. Однако на этот раз, отсутствующие электроны у пары хлорофилла А заменены электронами, перемещающимися из фотосистемы II через электронную транспортную цепь. Из молекулы носителя, пластохинона, к белку, цитохромному комплексу, и к другому белку, пластоцианину.Энергия, высвобождаемая в результате этого процесса, используется для закачивания протонов в тилакоидное пространство через цитохромный комплекс, создавая градиент протонов, который генерирует ATФ через хемиосмос. После прохождения электрона через этот центр реакции, он входит во вторую электронную транспортную цепь, начиная с белкового комплекса ферредоксина. Отсюда принимается одиночный электрон с помощью NADP+редуктазы, наряду с другими электроном и протоном, которые в конечном счёте формируют энергетический проект NADPH.

<ol>
	<li>Кардона, Т., Шао, С., Никсон,.Д. Усиление фотосинтеза в растениях: световые реакции. Очерки по биохимии. 62 (1), 85-94 (2018).</li>
</ol>

Фотосистема i

<h4>Обзор</h4>

Как и Фотосистема II (PS II), Фотосистема I (PS I) фиксирует фотоны и транспортируется через молекулы хлорофилла в центр реакции. В PS I, фотоны активизируют электроны, которые вошли PS I от PS II. Из центра реакции электрон высокой энергии отправляется через электронную транспортную цепочку и в конечном итоге соединяется с дополнительным электроном и протоном, чтобы уменьшить NADPв NADPH. Таким образом, подобно PS II, который захватывает энергию для генерации АТФ, PS I захватывает энергию для создания NADPH.

<h4>Фотосистема I</h4>

Пигменты светособирающего комплекса в Фотосистеме I поглощают фотоны и передают энергию реакционному центру (P700). После окисления электрон высокой энергии передается от специализированной пары хлорофилла а к первичному акцептору электронов. Однако на этот раз отсутствующие электроны из пары хлорофилла а заменяются электронами, движущимися из Фотосистемы II (вместо расщепления воды, как в ФС II). На своем пути от ФС II к ФС I электроны проходят через цепь переноса электронов, состоящую из молекулы-носителя пластохинона, двухбелкового цитохромного комплекса и пластоцианина.

После того, как электрон был возбужден в центре реакции PS I, он входит во вторую электронную транспортную цепь - белковый комплекс ферредоксин. Один электрон затем соединяется с другим электроном и протоном(H) , которые все принимаются NADPи редуктазой для формирования продукта NADPH. Таким образом, PS II улавливает энергию для создания протонного градиента, который в конечном счете используется для производства АТФ, а энергия, захваченная в PSI, используется для производства NADPH.

Фотосистема I

Несмотря на структурное сходство с фотосистемой II (PSII), фотосистема I (PSI) имеет другого поставщика электронов и акцептор электронов.
И то, и другое...

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photosystem i and nadph formation

Photosystem I helps convert light energy into NADPH during photosynthesis. It works with photosystem II in the thylakoid membrane of the chloroplast, but it uses a different electron supplier and a different electron acceptor.

Electrons excited in photosystem II move to photosystem I through an electron transport chain. That chain includes plastoquinone, the dual-protein cytochrome complex, and plastocyanin. As the electrons travel between the two photosystems, they lose energy and must be re-energized by photosystem I.

To do this, the pigments in the photosystem I antenna complex absorb another photon. The energy is passed to the reaction center called P700. P700 then sends a high-energy electron to NADP+ and forms NADPH.

Photosystem II uses light energy to build a proton gradient and make ATP. Photosystem I captures light energy to reduce NADP+ into NADPH. After ATP and NADPH are made, the cell has the energy it needs to build carbohydrate molecules for long-term storage in the stroma of the chloroplast.

Photosystem I and NADPH Formation

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Photosystem II is a multi-protein complex that captures light energy and begins the flow of electrons in photosynthesis. It uses bound pigments to harvest photons and send that energy to its reaction center. From there, the energy continues through the electron transport chain and on to photosystem I. The pigments in photosystem II include chlorophyll a, chlorophyll b, and carotenoids. They are arranged in two parts of the photosystem: the antenna complex and the reaction center. The antenna...

photosystem i

Photosystem I helps convert light energy into NADPH during photosynthesis. It works with photosystem II in the thylakoid membrane of the chloroplast, but it uses a different electron supplier and a different electron acceptor. Electrons excited in photosystem II move to photosystem I through an electron transport chain. That chain includes plastoquinone, the dual-protein cytochrome complex, and plastocyanin. As the electrons travel between the two photosystems, they lose energy and must be...

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key terms and concepts

Learn how photosystem I re-energizes electrons, reduces NADP+ to NADPH, and works with photosystem II to help build sugars in chloroplasts.

Photosystem I and NADPH Formation | JoVE K12

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Photosystem I and NADPH Formation

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