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The vestibular system detects the position and movement of the head through the labyrinth of the inner ear, which has three semicircular canals and two otolith organs.
The otolith organs contain vestibular hair cells that sit below the otolithic membrane, which is impregnated with otoconia, calcium carbonate crystals. When the head moves in certain ways, such as tilting forward this relatively heavy membrane shifts displacing the stereocilia on top of the hair cells generating a neural signal.
The semicircular canals also have vestibular hair cells, but their cilia are surrounded by a gelatinous cupula. Head movements such as rotations distort the cupula, moving the cilia.
For all vestibular hair cells, displacement towards the tallest cilium increases neurotransmitter release, while displacement in the opposite direction decreases it.
This information is then sent through the vestibular nerve to lower areas of the brain, such as the brain stem and cerebellum, so that gaze and balance can be rapidly adjusted to compensate for head movements. Some vestibular information travels further to the cerebral cortex, where it aids in the perception of body orientation.
The vestibular system is a set of inner ear structures that provide a sense of balance and spatial orientation. This system is comprised of structures…
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