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32.5:

์œ ์ „์ž ํ๋ฆ„

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Gene Flow

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์ง„ํ™”๋Š” ์‹œ๊ฐ„์— ์ง€๋‚˜๋ฉด์„œ ๋ชจ์ง‘๋‹จ ๋‚ด์—์„œ ์œ ์ „์  ๋ณ€ํ˜•์ด์žˆ์„ ๋•Œ๋งŒ ์ผ์–ด๋‚œ๋‹ค์ด ๋ณ€ํ™”์— ๊ธฐ์—ฌํ•˜๋Š” ํ•œ ๊ธฐ์ œ๊ฐ€์œ ์ „์ž ํ๋ฆ„, ์ฆ‰ ์œ ์ „์  ๋ณ€ํ˜•์ฒด๊ฐ€ ๋ชจ์ง‘๋‹จ์—์„œ์ด๋™ํ•˜๋Š” ๊ฒƒ์ด๋‹ค์œ ์ „์ž ํ๋ฆ„์€ ์œ ๊ธฐ์ฒด๊ฐ€ ํ•œ ๋ชจ์ง‘๋‹จ์—์„œ ๋‹ค๋ฅธ ๋ชจ์ง‘๋‹จ์œผ๋กœ ์ด๋™ํ•  ๋•Œ์ผ์–ด๋‚œ๋‹ค์–ด๋–ค ์ข…์—์„œ๋Š” ์ƒ์‹ ์„ธํฌ๋งŒ์ด๋ชจ์ง‘๋‹จ ์‚ฌ์ด์—์„œ ์ด๋™ํ•˜๋Š”๋ฐ ์‹๋ฌผ์˜ ๊ฝƒ๊ฐ€๋ฃจ๊ฐ€ ๋ฐ”๋žŒ์ด๋‚˜๋™๋ฌผ์— ์˜ํ•ด ์ด๋™ํ•œ๋‹ค์œ ์ „์ž ํ๋ฆ„์€ ์ƒˆ๋กœ์šด ์œ ์ „์  ๋ณ€์ด์ฒด๋ฅผ ๋„์ž…ํ•  ์ˆ˜ ์žˆ๋Š”๋ฐ, ์ด๋Š”๋ชจ์ง‘๋‹จ์—์„œ ๊ธฐ์กด ์œ ์ „ ๋ณ€์ด์ฒด์˜ ๋นˆ๋„๋ฅผ ๋ฐ”๊ฟ€ ์ˆ˜ ์žˆ๋‹ค์ž์—ฐ ์„ ํƒ์€ ์ƒ์กด๊ณผ ์ƒ์‹์„ ๋†’์ด๋Š” ํŠน์„ฑ์„ ์ง€์†์ ์œผ๋กœํ–ฅ์ƒ์‹œํ‚ค๋Š”๋ฐ ๊ทธ์™€๋Š” ๋‹ฌ๋ฆฌgene flow can either ์œ ์ „์ž ํ๋ฆ„์€ ์ ์‘ ํŠน์„ฑ์˜ ๋ฐœ์ƒ์„๋†’์ด๊ฑฐ๋‚˜ ์ค„์ผ ์ˆ˜ ์žˆ๋‹ค์ด๋ฅผํ…Œ๋ฉด, ์ด์ฃผํ•˜๋Š” ๋ชจ๊ธฐ๋Š”์‚ด์ถฉ์ œ์— ์ €ํ•ญํ•˜๋Š” ์œ ์ „ ๋ณ€์ด์ฒด ํ˜น์€ ๋Œ€๋ฆฝ ํ˜•์งˆ์„ํผ๋œจ๋ฆฐ๋‹ค๊ทธ๋ž˜์„œ, ์ƒˆ๋กœ์šด ๋Œ€๋ฆฝ ํ˜•์งˆ์„ ๊ฐ€์ง„ ๋ชจ๊ธฐ๋Š”๋” ์„ฑ๊ณต์ ์œผ๋กœ ์ƒ์กดํ•˜๊ณ  ์ƒ์‹ํ•œ๋‹ค๋ฐ˜๋ฉด์— ๋ฐ•์ƒˆ์˜๋ธ”๋ฆฌ๋žœ๋“œ ์„ฌ์œผ๋กœ์˜ ์ด์ฃผ๋Š”์œ ์ „์ž ํ๋ฆ„์ด ์–ด๋–ป๊ฒŒ ํ•ด๋กœ์šธ ์ˆ˜ ์žˆ๋Š”์ง€ ๋ณด์—ฌ์ค€๋‹ค๋ณธํ† ์—์„œ ์ด์ฃผํ•˜๋Š” ์ƒˆ๋Š”๋‚ณ๋Š” ์•Œ ๊ฐฏ์ˆ˜๊ฐ€ ๋” ํฐ ์œ ์ „์ž๋ฅผ ๊ฐ–๊ณ  ์žˆ๋Š”๋ฐ์„ฌ์˜ ์ƒˆ๋“ค์€ ์•Œ ๊ฐฏ์ˆ˜๊ฐ€ ๋” ์ ๋‹ค๋ธ”๋ฆฌ๋žœ๋“œ์˜ ์ค‘์•™์—๋Š” ์ด์ฃผ๊ฐ€ ์„ธ ๋ฐฐ๋‚˜ ๋†’์€๋ฐ๋ฐ•์ƒˆ ๋ชจ์ง‘๋‹จ์ด ๋™๋ถ€ ๋ธ”๋ฆฌ๋žœ๋“œ๋ณด๋‹ค๋‚ฎ์€ ์ƒ์กด์œจ์„ ๊ฐ–๊ณ  ์žˆ๊ณ ์ด์ฃผ์œจ๋„ ์ ๋‹ค์‹œ๊ฐ„์ด ์ง€๋‚˜๋ฉด์„œ, ์œ ์ „์ž ํ๋ฆ„์€ ๋ชจ์ง‘๋‹จ ์‚ฌ์ด์˜ ์œ ์ „์  ์ฐจ์ด๋ฅผ์ค„์ด๊ณ , ๊ฐ€๋” ๊ณตํ†ต ์œ ์ „์ž ํ’€๋กœ ์ด์–ด์ง„๋‹คํ•˜์ง€๋งŒ, ์œ ์ „์ž ํ๋ฆ„์ด ์ œํ•œ๋  ๋•Œ๋Š” ์œ ์ „์  ์ฐจ์ด๊ฐ€์‹œ๊ฐ„์ด ์ง€๋‚˜๋ฉด์„œ ์ฆํญ๋œ๋‹ค์ˆ˜๋ฐฑ๋งŒ ๋…„ ์ „์— ์ค„์–ด๋“œ๋Š” ๋Œ€์–‘ ์ˆ˜์œ„๋กœํŒŒ๋‚˜๋งˆ ํ•ดํ˜‘์ด ๋งŒ๋“ค์–ด์กŒ๊ณ  ์ด ํ•ดํ˜‘์€๋‚จ๋ถ ์•„๋ฉ”๋ฆฌ์นด๋ฅผ ์—ฐ๊ฒฐํ•œ๋‹ค๊ทธ ๊ฒฐ๊ณผ ์ด์ข… ๊ต๋ฐฐํ•˜๋Š” ํ•ด์–‘ ๋ชจ์ง‘๋‹จ์ด๋ถ„๋ฆฌ๋˜๊ณ  ๋ง์•˜๋‹ค๊ทธ ํ•ดํ˜‘์˜ ์–‘ ์ชฝ ๋ชจ์ง‘๋‹จ์€์Šค์Šค๋กœ ๋ฒˆ์‹์„ ํ–ˆ์ง€๋งŒ ์„œ๋กœ๋Š”ํ•˜์ง€ ๋ชปํ–ˆ๋‹ค๋šœ๋ ทํ•˜๊ณ  ๋ฌด์ž‘์œ„ํ•œ ๋ณ€์ด๊ฐ€ ๋ถ„๋ฆฌ๋œ ๋ชจ์ง‘๋‹จ์—์„œ ์ƒ๊ฒจ๋‚˜๊ณ ํ™˜๊ฒฝ ์กฐ๊ฑด ๋•Œ๋ฌธ์— ์—ฌ๋Ÿฌ ํŠน์ง•์ด์ž์—ฐ ์„ ํƒ๋˜์—ˆ๋‹ค๊ฒฐ๊ตญ, ์œ ์ „์ € ์ฐจ์ด๊ฐ€ ์ƒ์‹ ํŠน์„ฑ์—์„œ๋‚˜ํƒ€๋‚˜๊ณ , ๋šœ๋ ทํ•œ ์ข…์ด ํƒ„์ƒํ–ˆ๋Š”๋ฐ๊ฐ€๊นŒ์ด ์žˆ์Œ์—๋„ ๋ถˆ๊ตฌํ•˜๊ณ  ๋” ์ด์ƒ ๊ต๋ฐฐ๋  ์ˆ˜ ์—†์—ˆ๋‹ค๊ทธ๋ž˜์„œ, ์œ ์ „์ž ํ๋ฆ„์€ ๋ง‰๊ฐ•ํ•œ ์ง„ํ™” ๊ธฐ์ œ๋กœ์„œ๊ฐœ๋ณ„ ๋ชจ์ง‘๋‹จ์˜ ์œ ์ „์  ๋‹ค์–‘์„ฑ์„ ํ–ฅ์ƒ์‹œํ‚ค๊ณ ๊ทธ๋“ค ์‚ฌ์ด์˜ ์œ ์ „์  ์ฐจ์ด๋ฅผ์ œํ•œํ•œ๋‹ค

32.5:

์œ ์ „์ž ํ๋ฆ„

์œ ์ „์ž ํ๋ฆ„(gene flow)์€ ๋ฐฐ์šฐ์ž(gamete)์˜ ํ™•์‚ฐ ๋˜๋Š” ๊ฐœ์ฒด์˜ ์ด์ฃผ(migration)๋ฅผ ํ†ตํ•ด ์œ ์ „์ž๊ฐ€ ํ•œ ์ง‘๋‹จ์—์„œ ๋‹ค๋ฅธ ์ง‘๋‹จ์œผ๋กœ ์ด๋™ํ•˜๋Š” ๊ฒƒ์ž…๋‹ˆ๋‹ค.

์ด ํ˜„์ƒ์€ ๋ชจ๋“  ์œ ๊ธฐ์ฒด์—์„œ ์ค‘์š”ํ•œ ์ง„ํ™”์  ์—ญํ• ์„ ํ•˜๋ฉฐ, ์œ ์ „์ž ํ๋ฆ„ ์†๋„์— ๋”ฐ๋ผ ์ง‘๋‹จ๋“ค ์‚ฌ์ด์—์„œ ์œ ์ „์  ๋‹ค์–‘์„ฑ(genetic diversity)์„ ์œ ๋„ํ•˜๊ฑฐ๋‚˜ ์œ ์ „์  ๋“ฑ์งˆ์„ฑ(genetic homogeneity)์„ ๋ฐœ์ƒ์‹œํ‚ต๋‹ˆ๋‹ค. ์œ ์ „์ž ํ๋ฆ„ ์†๋„๊ฐ€ ๋Š๋ฆด ๋•Œ ์ง‘๋‹จ์— ์ƒˆ๋กœ์šด ๋Œ€๋ฆฝ์œ ์ „์ž(allele)๊ฐ€ ๋„์ž…๋˜๋ฉด ์œ ์ „์  ๋‹ค์–‘์„ฑ์ด ์ƒ๊น๋‹ˆ๋‹ค. ๋ฐ˜๋ฉด์—, ์œ ์ „์ž ํ๋ฆ„ ์†๋„๊ฐ€ ๋น ๋ฅผ ๋• ์ง‘๋‹จ ์‚ฌ์ด์˜ ์œ ์ „์  ๋ณ€ํ™”๋ฅผ ๊ฐ์†Œ ์‹œ์ผœ ๋“ฑ์งˆ์„ฑ์„ ์ฆ๊ฐ€์‹œํ‚ต๋‹ˆ๋‹ค.

์ด๋™์„ฑ(mobility)์€ ์ง‘๋‹จ ๊ฐ„์˜ ์œ ์ „์ž ํ๋ฆ„ ์†๋„์— ์˜ํ–ฅ์„ ๋ฏธ์น˜๋Š” ์ค‘์š”ํ•œ ์š”์†Œ์ž…๋‹ˆ๋‹ค. ์ด๋™์„ฑ์ด ๋†’์€ ์œ ๊ธฐ์ฒด๋Š” ์ด์ฃผ๋ฅผ ํ†ตํ•ด ์œ ์ „์ž ํ๋ฆ„์— ๊ธฐ์—ฌํ•  ๊ฐ€๋Šฅ์„ฑ์ด ๋” ํฝ๋‹ˆ๋‹ค. ์‹๋ฌผ๊ณผ ๊ฐ™์€ ์ •์ฃผํ˜• ์œ ๊ธฐ์ฒด๋Š” ๋™๋ฌผ์ด๋‚˜ ๋ฐ”๋žŒ์— ์˜ํ•ด ๋จผ ๊ณณ์œผ๋กœ ์˜ฎ๊ฒจ์ง€๋Š” ๊ฝƒ๊ฐ€๋ฃจ์™€ ์”จ์•—์„ ํ†ตํ•ด ์œ ์ „์ž ํ๋ฆ„์„ ์ด‰์ง„ํ•  ์ˆ˜ ์žˆ์Šต๋‹ˆ๋‹ค. ๊ทธ๋Ÿฌ๋‚˜ ์ด์ฃผ ์ž์ฒด๊ฐ€ ํ•ญ์ƒ ์œ ์ „์ž์˜ ํ๋ฆ„์„ ๋ณด์žฅํ•˜๋Š” ๊ฒƒ์€ ์•„๋‹™๋‹ˆ๋‹ค. ์ด์ฃผ๋ฅผ ํ†ตํ•œ ์œ ์ „์ž ํ๋ฆ„์€ ๊ฑฐ์ฃผ์ž์™€ ์ด์ฃผ์ž ์‚ฌ์ด์˜ ์œ ์ „์ž ๋˜๋Š” ์œ ์ „ ๋ฌผ์งˆ์˜ ๊ตํ™˜์ด ๋™๋ฐ˜๋˜์–ด์•ผ ํ•ฉ๋‹ˆ๋‹ค.

๋ฌผ๋ฆฌ์ , ์ƒ์‹์  ์žฅ๋ฒฝ์€ ์œ ์ „์ž ํ๋ฆ„์„ ๋ฐฉํ•ดํ•  ์ˆ˜ ์žˆ์Šต๋‹ˆ๋‹ค. ์˜ˆ๋ฅผ ๋“ค์–ด ์ด์†Œ์  ์ข…๋ถ„ํ™”(allopatric speciation)๋Š” ์ง€๋ฆฌ์  ์žฅ๋ฒฝ์ด ๋™์ผํ•œ ์ข…์˜ ์ง‘๋‹จ์„ ๊ฒฉ๋ฆฌํ•˜์—ฌ ์œ ์ „์ž ํ๋ฆ„์„ ์ œํ•œํ•  ๋•Œ ๋ฐœ์ƒํ•ฉ๋‹ˆ๋‹ค. ๋งŒ์•ฝ ์ง‘๋‹จ์ด ์ถฉ๋ถ„ํžˆ ๋ถ„๋ฆฌ๋˜์–ด ์žฅ๋ฒฝ์ด ์ œ๊ฑฐ๋˜์—ˆ์„ ๋•Œ ๋‹ค๋ฅธ ์ง‘๋‹จ๊ณผ ํ•จ๊ป˜ ๋” ์ด์ƒ ์ƒ์กด ๊ฐ€๋Šฅํ•œ ์ž์†์„ ๋‚ณ์„ ์ˆ˜ ์—†๋‹ค๋ฉด, ๋‘ ์ง‘๋‹จ์€ ๋ณ„๋„์˜ ์ข…์œผ๋กœ ๋ถ„๋ฅ˜๋  ์ˆ˜ ์žˆ์Šต๋‹ˆ๋‹ค.

์ธ๊ฐ„์ด ๋ณด์กฐํ•˜๋Š” ์œ ์ „์ž ํ๋ฆ„์€ ์œ ์ „์ž ๊ตฌ์กฐ(genetic rescue)์— ๋„์›€์„ ์ค„ ์ˆ˜ ์žˆ์Šต๋‹ˆ๋‹ค. ์†Œ๊ทœ๋ชจ ์ง‘๋‹จ์—์„œ์˜ ๋†’์€ ๋™๊ณ„๊ต๋ฐฐ(inbreeding)๋ฅ ์€ ๋‹ค์–‘์„ฑ์„ ๋‚ฎ์ถ”๊ณ , ์ ์‘๋„(fitness)๋ฅผ ๊ฐ์†Œ์‹œํ‚ค๋ฉฐ, ๋ฉธ์ข… ์œ„ํ—˜์„ ์ฆ๊ฐ€์‹œํ‚ต๋‹ˆ๋‹ค. ์ธ๊ฐ„์˜ ๊ฐ„์„ญ์„ ํ†ตํ•ด ํ˜ˆ์—ฐ์ด ์•„๋‹Œ ๊ฐœ์ฒด๋‚˜ ์œ ๊ธฐ์ฒด์˜ ๋„์ž…์€ ๋™๊ณ„๊ต๋ฐฐ ๋ฒ”์œ„๋ฅผ ์ค„์ด๊ณ  ๋‹ค์–‘์„ฑ์„ ๊ฐœ์„ ํ•˜๋ฉฐ ์ „๋ฐ˜์ ์ธ ์ ์‘๋„๋ฅผ ํ–ฅ์ƒํ•  ์ˆ˜ ์žˆ์Šต๋‹ˆ๋‹ค.

Suggested Reading

McDonald, Bruce A. "Population Genetics of Plant Pathogens." Population Genetics of Plant Pathogens. The American Phytopathological Society (APS), n.d. Accessed February 4, 2020. [Source]

Morjan, Carrie L., and Loren H. Rieseberg. "How Species Evolve Collectively: Implications of Gene Flow and Selection for the Spread of Advantageous Alleles." Molecular Ecology. 13, no. 6 (2004): 1341-356. [Source]