Denver Museum of Nature and Science 1 article published in JoVE Neuroscience Denver Papillae Protocol for Objective Analysis of Fungiform Papillae Tiffany M. Nuessle1, Nicole L. Garneau1, Meghan M. Sloan1, Stephanie A. Santorico2 1Health Sciences Department, Denver Museum of Nature & Science, 2Department of Mathematical and Statistical Sciences, University of Colorado Denver Here we present a protocol to measure fungiform papilla density from digital photographs. This method builds prioritization and objective characteristic metrics into the original descriptive work on fungiform papillae by Miller & Reedy (1990).